Publications

Matsumura, Y., Lima, S.P., Rafael, J.A., Câmara, J.T., Beutel, R.G., Gorb, S.N., 2022. Distal leg structures of Zoraptera – did the loss of adhesive devices curb the chance of diversification? Arthropod Struct. Dev., 68: 101164. https://doi.org/10.1016/j.asd.2022.101164External link

Yavorskaya, M.I., Jałoszyński, P., Beutel, R.G., 2022. A unique case of commensalism: The beaver beetle Platypsyllus castoris (Leiodidae, Coleoptera) and its morphological adaptations. J. Morphol., 284(1): e21532. https://doi.org/10.1002/jmor.21532External link

Luo, X.Z., Jałoszyński, P., Beutel, R.G., 2022. The thoracic anatomy of Pselaphus heisei (Pselaphinae, Staphylinidae, Coleoptera). J. Morphol., 284(1): e21527. https://doi.org/10.1002/jmor.21527External link

Fikáček, M., Yamamoto, S., Matsumoto, K., Beutel, R.G., David R. Maddison, D.R., 2022. Phylogeny and systematics of Sphaeriusidae (Coleoptera: Myxophaga): minute living fossils with underestimated past and present-day diversity. Syst. Entomol.: 1–17. https://doi.org/10.1111/syen.12571External link

Boudinot, B.E., Fikáček, M., Lieberman, Z.E., Kusy, D., Bocak, L., McKenna, D.D., Beutel, R.G., 2022. Systematic bias and the phylogeny of Coleoptera — a response to Cai et al. (2022) following the responses to Cai et al. (2020). Syst. Entomol., 2022, 1–10. https://doi.org/10.1111/syen.12570External link

Cho, H.-W., Kwon, M.Ch., Kim, S.K., Beutel, R.G., 2022. Morphology of larvae and pupae of the genus Autocrates (Coleoptera: Trictenotomidae) and its phylogenetic implications. Acta Entomol. Mus. Natl. Pragae 62(1): 197–210. https://doi.org/10.37520/aemnp.2022.015External link

Tozetto, L., Chaul, J.C.M., Boudinot, B.E., Lattke, J.E., 2022. Review of the Leptogenys unistimulosa species group (Hymenoptera: Formicidae) with the description of a new Amazonian species. Rev. Bras. Entomol., 66(3), e20220045. https://doi.org/10.1590/1806-9665-RBENT-2022-0045External link

Benda, D., Pohl, H., Beutel, R., Straka, J., 2022. Two new species of Xenos (Strepsiptera: Xenidae), parasites of social wasps of the genus Mischocyttarus (Hymenoptera: Vespidae) in the New World. Acta Entomol. Mus. Natl. Pragae 62(1), 185–195. https://doi.org/10.37520/aemnp.2022.014External link

Richter, A., Boudinot, B.E., Yamamoto, S., Katzke, J., Beutel, R.G., 2022. The first reconstruction of the head anatomy of a Cretaceous insect, †Gerontoformica gracilis (Hymenoptera, Formicidae), and the early evolution of the ants. Insect Syst. Divers., 6(5), 4, 1–80. https://doi.org/10.1093/isd/ixac013External link

Boudinot, B.E., Richter, A.K., Hammel, J., Szwedo, J., Bojarski, B., Perrichot, V., 2022. Genomic-phenomic reciprocal illumination: †Desyopone gen. nov., an exceptional aneuretine-like fossil ant from Ethiopian amber (Hymenoptera: Formicidae: Ponerinae). Insects, 13, 796. https://doi.org/10.3390/insects13090796External link

Jandausch, K., Michels, J., Kovalev, A., Gorb, S.N., van de Kamp, T., Beutel, R.G., Niehuis, O., Pohl, H., 2022. Have female twisted-wing parasites (Insecta: Strepsiptera) evolved tolerance traits as response to traumatic penetration? PeerJ 10, e13655. https://doi.org/10.7717/peerj.13655External link

Li, D., Friedrich, F., Jandausch, K., Pohl, H., Liu, X., Beutel, R.G., 2022. Unearthing underground predators: The head morphology of larvae of the moth lacewing genus Ithone Newman (Neuroptera: Ithonidae) and its functional and phylogenetic implications. Syst. Entomol., 47(4), 618–636. https://doi.org/10.1111/syen.12556External link

Boudinot, B.E., Yan, E.V. Prokop, J., Luo, X.-Z., Beutel, R.G., 2022. Permian parallelisms: Reanalysis of †Tshekardocoleidae sheds light on the earliest evolution of the Coleoptera. Syst. Entomol., XX, 1–28. https://doi.org/10.1111/syen.12562External link

Perrichot, V., Boudinot, B.E., Engel, M.S., Xu, C., Bojarski, B., Szwedo, J., 2022. Ants (Hymenoptera: Formicidae) in Miocene Ethiopian amber: filling gaps in the geological record of African terrestrial biota. Zool. J. Linn. Soc., 1–17. https://doi.org/10.1093/zoolinnean/zlac053External link

Lieberman, Z., Billen, J., van de Kamp, T., Boudinot, B.E., 2022. The abdominal anatomy of the worker of Amblyopone australis ants (Formicidae: Amblyoponinae), with special reference to skeletomusculature. J. Morphol. https://doi.org/10.1002/jmor.21471External link

Aibekova, L., Boudinot, B.E., Beutel, R.G., Richter, A., Keller, R.A., Hita-Garcia, F., Economo, E.P., 2022. The skeletomuscular system of the mesosoma of Formica rufa workers (Hymenoptera: Formicidae). Insect. Syst. Divers. 6, 1–26. https://doi.org/10.1093/isd/ixac002External link

Benda, D., Pohl, H., Nakase, Y., Beutel, R., Straka, J., 2022. A generic classification of Xenidae (Strepsiptera) based on the morphology of the female cephalothorax and male cephalotheca with a preliminary checklist of species. ZooKeys 1093, 1–134. https://doi.org/10.3897/zookeys.1093.72339External link

Beutel, R.G., Wipfler, B., Meusemann, K., Pohl, H., 2022. Insect systematics from Hennig to transcriptomes – anatomy and phylogenomics converge upon a robust phylogeny. Atti Accad. Naz. Ital. Entomol. 69, 93–99.

Boudinot, B.E., Borowiec, M.L., Prebus, M.M. 2022. Phylogeny, evolution, and classification of the ant genus Lasius, the tribe Lasiini, and the subfamily Formicinae (Hymenoptera: Formicidae). Syst. Entomol. 47, 113–151. https://doi.org/10.1111/syen.12522External link

Boudinot, B.E., Richter, A., Beutel, R.G., Katzke, J., Keller, R.A., Economo, E.P., Yamamoto, E., 2022. Evidence for the evolution of eusociality in stem ants and a systematic revision of †Gerontoformica (Hymenoptera, Formicidae). Zool. J. Linn. Soc. https://doi.org/10.1093/zoolinnean/zlab097External link

Jałoszyński, P., Luo, X.-Zh., Beutel, R.G., 2022. Evolution of cephalic structures in extreme myrmecophiles: a lesson from Clavigeritae (Coleoptera: Staphylinidae: Pselaphinae). Cladistics. https://doi.org/10.1111/cla.12498External link

Möckel, L., Meusemann, K., Misof, B., Schwartze, V.U., Fine Licht, H.H., Voigt, K., Stielow, B., de Hoog, S., Beutel, R.G., Buellesbach, J., 2022. Phylogenetic revision and patterns of host specificity in the fungal subphylum Entomophthoromycotina. Microorganisms 10, 256, https://doi.org/10.3390/microorganisms10020256External link.

Niermann, J., Brehm, G., 2022. The number of sampled moths is more affected by microhabitat choice than by UV lamp type in a grassland habitat. Eur. J. Entomol. 119, 36–42. https://doi.org/10.14411/eje.2022.004External link

Bao, T, Wedmann, S., Grímsson, F. Beutel, R.G., Seyfullah, L., Bao, L., Jarzembowski, E.A., 2021. Was the kateretid beetle Pelretes really a Cretaceous angiosperm pollinator? Nature Plants 8, 38–40. https://doi.org/10.1038/s41477-021-00893-2

Beutel, R.G., Friedrich, F., Economo, E.P., 2021. Patterns of morphological simplification and innovation in the megadiverse Holometabola (Insecta). Cladistics (early view). https://doi.org/10.1111/cla.12483

Beutel, R.G., Luo, X.-Zh., Yavorskaya, M.I., Jałoszyński, P., 2021. Structural megadiversity in leaf litter predators-the head anatomy of Pselaphus heisei (Pselaphinae, Staphylinidae, Coleoptera). Arthropod Syst. Phyl. 79, 443.

Boudinot, B.E., Beutel, R.G., Gorb, S.N., Polilov, A.A., 2021. Functional diversity of attachment and grooming leg structures is retained in all but the smallest insects. J. Zool. 313(2), 99–113.

Boudinot, B.E., Moosdorf, O.T.D., Beutel, R.G., Richter, A., 2021. Anatomy and evolution of the head of Dorylus helvolus (Formicidae: Dorylinae): Patterns of sex- and caste-limited traits in the sausagefly and the driver ant. J. Morphol. 282, 1616–1658. Doi: 10.1002/jmor.21410.

Brehm, G., Niermann, J., Jaimes Nino, L.M., Enseling, D., Jüstel, T., Fiedler K. 2021. Moths are strongly attracted to ultraviolet and blue radiation. Insect Conserv. Diversity 14, 188–198

Enkhtur, K., Brehm, G., Boldgiv, B., Pfeiffer, M., 2021. Alpha and beta diversity of moths along a latitudinal gradient in the Eastern Palearctic. Sci. Rep. 11, 15018.

Enkhtur, K., Brehm, G., Boldgiv, B., Pfeiffer, M., 2021. Effects of grazing on macro-moth assemblages in two different biomes in Mongolia. Ecol. Indic. 133, 108421.

Fiedler, K., Brehm, G., 2021. Aposematic Coloration of Moths Decreases Strongly along an Elevational Gradient in the Andes. Insects 12, 903.

Fischer, S., Laue, M., Müller, C.H.G., Meinertzhagen, I.A., Pohl, H., 2021. Ultrastructural 3D re-construction of the smallest known insect photoreceptors: The stemmata of a first instar larva of Strepsiptera (Hexapoda). Arthr. Struct. Develop 62, 101055. doi: 10.1016/ j.asd.2021.101055

Gaona, F.P., Iniguez-Armijos, C., Brehm, G., Fiedler, K., Espinosa, C.I., 2021. Drastic loss of insects (Lepidoptera: Geometridae) in urban landscapes in a tropical biodiversity hotspot. J. Insect Conserv. 1–11.

Luo, C.H., Beutel, R.G., Thomson, U.R., Zheng, D.R., Li, J. H., Zhao, X.Y., Zhang, H.-Ch., Wang, B., 2021. Beetle or roach: systematic position of the enigmatic Umenocoleidae based on new material from Zhonggou Formation in Jiuquan, Northwest China, and a morphocladistic analysis. Palaeoworld (in press).

Luo, X.Z., Hlaváč, P., Jałoszyński, P. & Beutel, R.G., 2021. In the twilight zone – the head morphology of Bergrothia saulcyi (Pselaphinae, Staphylinidae, Coleoptera), a beetle with adaptations to endogean life but living in leaf litter. J. Morph., online early.

Luo, X.Z., Jałoszyński, P., Stoessel, A., Beutel, R.G., 2021. The specialized thoracic skeletomuscular system of the myrmecophile Claviger testaceus (Pselaphinae, Staphylinidae, Coleoptera). Org. Div. Evol. 21, 317–335.

Murillo-Ramos, L., Friedrich, E., Williams, S., Wahlberg, N., Brehm, G., Sihvonen, P., 2021. A morphological appraisal of the new subfamily Epidesmiinae (Lepidoptera: Geometridae) with an overview of all geometrid subfamilies. Zool. J. Linn. Soc.

Murillo-Ramos, L., Sihvonen, P., Brehm, G., Ríos-Malaver, I.C., Wahlberg, N., 2021. A database and checklist of geometrid moths (Lepidoptera) from Colombia. Biodivers. data J. 9.

Pohl, H., 2021. Die Stammgruppe der Fächerflügler (Insecta, Strepsiptera). Archiv NLk. M-V. 58, 58–70. doi:10.30819/anlk.58.06

Pohl, H., Wipfler, B., Boudinot, B., Beutel, R.G., 2021. On the value of Burmese amber for understanding insect evolution: Insights from †Heterobathmilla – an exceptional stem group genus of Strepsiptera (Insecta). Cladistics, 37(2), 211–229.

Qvarnström, M., Fikáček, M., Wernström, J.V., Huld, S., Beutel, R.G., Arriaga-Varela, E., Ahlberg, P.E., Niedźwiedzki, G. 2021. Exceptionally preserved beetles in a Triassic coprolite of putative dinosauriform origin. Curr, Biol. 31(15), 3374–3381.

Richter, A., Keller, R.A., Hita Garcia, F., Billen, J., Katzke, J., Boudinot, B.E., Economo, E.P., Beutel, R.G., 2021. Head anatomy of Protanilla lini (Leptanillinae, Formicidae, Hymenoptera) and a hypothesis of their mandibular movement. Myrmecol. News 31, 107–136. Doi: 10.25849/myrmecol.news_031:107.

Vasilikopoulos, A., Balke, M., Kukowka, S., Pflug, J.M., Martin, S., Meusemann, K., Hendrich, L., Mayer, Ch., Maddison, D.R., Niehuis, O., Beutel, R.G., Misof, B., 2021. Phylogenomic analyses clarify the pattern of evolution of Adephaga (Coleoptera) and highlight phylogenetic artifacts due to model misspecification and excessive data trimming. Syst. Entomol. 46(4), 991–1018.

Beutel, R. G., Richter, A., Keller, R. A., Hita Garcia, F., Matsumura, Y., Economo, E. P., Gorb, S.N., 2020. Distal leg structures of the Aculeata (Hymenoptera): A comparative evolutionary study of Sceliphron (Sphecidae) and Formica (Formicidae). J. Morph. 281, 737–753.

Beutel, R.G., Liu, Z., Fikáček, M., Ren, D., Pang, H., Ślipiński, A., 2020. Burmapseudomorphus planus gen. et sp. nov. – a Late Cretaceous stem group member of the specialized Pseudomorphini (Carabidae, Coleoptera) from northern Myanmar. Cretac. Res. 107, 104274.

Beutel, R.G., Luo, X., Wipfler, B., 2020. Is †Umenocoleus a roach or a beetle (Dictyoptera or Coleoptera)? Palaeoentomol. 3(1), 96–102.

Boudinot, B.E., Beutel, R.G., Gorb, S.N., Polilov, A.A. 2020. Functional diversity of attachment and grooming leg structures is retained in all but the smallest insects. J. Zool. 313(2), 99–113.

Jałoszyński, P., Luo, X.Z. & Beutel, R.G. 2020. Profound head modifications in Claviger testaceus (Pselaphinae, Staphylinidae, Coleoptera) facilitate integration into communities of ants. Journal of Morphology 281(9): 1072–1085.

Jałoszyński, P., Luo, X.Z., Hammel, J.U., Yamamoto, S., Beutel, R.G., 2020. The mid-Cretaceous †Lepiceratusgen. nov. and the evolution of the relict beetle family Lepiceridae (Insecta: Coleoptera: Myxophaga). J. Syst. Palaeont., 1–14.

Matsumura, Y., Beutel, R.G., Rafael, J.A., Yao, I., Câmara, J.T., Lima, Sh.P., Yoshizawa, K., 2020. The evolution of Zoraptera. Syst. Entomol. 45(2), 349–364.

Pohl, H., Gorb, E.V., Gorb, S.N., 2020. Traction force measurements on male Strepsiptera (Insecta) revealed higher forces on smooth compared with hairy substrates. Journal of Experimental Biology 223, 1–7. doi:10.1242/jeb.223784

Pohl, H., Hammel, J.U., Richter, A., Beutel, R.G., 2020. First fossil late instar larva of Strepsiptera, Photon Science – Highlights and Annual Report. Hamburg.

Rabl, D., Alonso-Rodriguez, A.M., Brehm, G., Fiedler, K., 2020. Trait variation in moths mirror small-scaled ecological gradients in a tropical landscape. Insects 1, 612.

Rabl, D., Gottsberger, B., Brehm, G., Hofhansl, F., Fiedler, K., 2020. Moth assemblages in Costa Rica rain forest mirror small-scale topographic heterogeneity. Biotropica 52, 288–301.

Richter, A., Garcia, F.H., Keller, R.A., Billen, J., Economo, E.P., Beutel, R.G., 2020. Comparative analysis of worker head anatomy of Formica and Brachyponera (Hymenoptera: Formicidae). Arthropod Syst. Phyl. 78(1), 133–170.

Sihvonen, P., Murillo-Ramos, L., Brehm, G., Staude, H., Wahlberg, N., 2020. Molecular phylogeny of Sterrhinae moths (Lepidoptera: Geometridae): towards a global classification. Syst. Entomol. 45, 606–634.

Tröger, D., Grabe, V., Beutel, R.G., Pohl, H. 2020. The endoparasitic larval stages of Eoxenos laboulbenei: an atypical holometabolan development (Strepsiptera, Mengenillidae). Arthropod Str. Dev. 56, 100932.

Vasilikopoulos, A., Gustafson, G.T., Balke, M., Niehuis, O., Beutel, R.G., Misof, B., 2020. Resolving the phylogenetic position of Hygrobiidae (Coleoptera: Adephaga) requires objective statistical tests and exhaustive phylogenetic methodology: a response to Cai et al. 2020. Mol. Phyl. Evol. 106923–106923.

Vasilikopoulos, A., Misof, B., Meusemann, K., Lieberz, D., Flouri, T., Beutel, R.G., Niehuis, O., Wappler, T., Rust, J., Peters, R.S., Donath, A., Podsiadlowski, L., Mayer, Ch., Bartel, D., Böhm, A., Liu1, Sh., Kapli, P., Greve, C., Jepson, J.E., Liu, X., Zhou, X., Aspöck, H., Aspöck, U., 2020. An integrative phylogenomic approach to elucidate the evolutionary history and divergence times of Neuropterida (Insecta: Holometabola). BMC Evolut. Biol. 20, 64.

Antunes‐Carvalho, C., Ribera, I., Beutel, R.G., Gnaspini, P., 2017. Morphology‐based phylogenetic reconstruction of Cholevinae (Coleoptera: Leiodidae): a new view on higher‐level relationships. Cladistics 35(1), 1–41.

Beutel, R.G., Fabian, B., Pohl, H., Friedrich, F., 2019. 2.4. Morphology of larvae. In: Beutel, R.G., Friedrich, F. (Eds.), Handbook of Zoology, Vol. IV Arthropoda: Insecta. Nannomecoptera and Neomecoptera. Walter De Gruyter, Berlin, New York.

Beutel, R.G., Jandausch, K., 2019. 3.3.9. Digestive system. In: Beutel, R.G., Friedrich, F. (Eds.), Handbook of Zoology, Vol. IV Arthropoda: Insecta. Nannomecoptera and Neomecoptera. Walter De Gruyter, Berlin, New York.

Brehm, G., Murillo-Ramos, L., Sihvonen, P., Hausmann, A., Schmidt, B.C., Õunap, E., Moser, A., Mörtter, R., Bolt, D., Bodner, F., others, 2019. New World geometrid moths (Lepidoptera: Geometridae): Molecular phylogeny, biogeography, taxonomic updates and description of 11 new tribes. Arthropod Syst. & Phylogeny.

Brehm, G., Zeuss, D., Colwell, R.K., 2019. Moth body size increases with elevation along a complete tropical elevational gradient for two hyperdiverse clades. Ecography (Cop.). 42, 632–642.

Jaimes Nino, L.M., Mörtter, R., Brehm, G., 2019. Diversity and trait patterns of moths at the edge of an Amazonian rainforest. J. Insect Conserv. 23, 751–763.

Krämer, J., Pohl, H., Predel, R., 2019. Venom collection and analysis in the pseudoscorpion Chelifer cancroides(Pseudoscorpiones: Cheliferidae). Toxicon 162, 15–23. doi:10.1016/j.toxi-con. 2019.02.009

Liu, S.-P., Richter, A., Stoessel, A., Beutel, R.G., 2019. The mesosomal anatomy of Myrmecia nigrocincta workers and evolutionary transformations in Formicidae (Hymenoptera). Arthropod Syst. Phyl. 77 (1), 1–19.

Luo, X.-Zh., Antunes‐Carvalho, C., Ribera, I. & Beutel, R.G., 2019. The thoracic morphology of the troglobiontic cholevine species Troglocharinus ferreri (Coleoptera, Leiodidae). Arthropod Str. Dev. 53, 100900.

Luo, X.Z., Antunes‐Carvalho, C., Wipfler, B., Ribera, I., Beutel, R.G., 2019. The cephalic morphology of the troglobiontic cholevine species Troglocharinus ferreri (Coleoptera, Leiodidae). J. Morph. 280(8), 1207–1221.

Mashimo, Y., Müller, P., Pohl, H., Beutel, R.G., 2019. Zorotypus pecten, a new species of Zoraptera (Insecta) from mid-Cretaceous Burmese amber. Zootaxa 4651(3), 565–577.

McKenna, D.D., Shin, S., Ahrens, D., Balke, M., Beza-Beza, C., Clarke, D.J., Donath, A., Escalona, H.E., Friedrich, F., Letsch, H., Liu, Sh., Maddison, D., Mayer, Ch., Misof, B., Murin, P.J., Niehuis, O., Peters, R.S., Podsiadlowski, L., Pohl, H., Scully, E.D., Yan, E.V., Zhou, X., Ślipiński, A., Beutel, R.G., 2019. The evolution and genomic basis of beetle diversity. PNAS 116(49), 24729–24737.

Murillo-Ramos, L., Brehm, G., Sihvonen, P., Hausmann, A., Holm, S., Ghanavi, H.R., Õunap, E., Truuverk, A., Staude, H., Friedrich, E., others, 2019. A comprehensive molecular phylogeny of Geometridae (Lepidoptera) with a focus on enigmatic small subfamilies. PeerJ 7, e7386.

Pohl, H., Beutel, R.G., 2019. Effects of miniaturization in primary larvae of Strepsiptera (Insecta). Arthropod Str. Dev. 48, 49–55.

Pohl, H., Hammel, J.U., Richter, A., Beutel, R.G., 2019. The first fossil free-living late instar larva of Strepsiptera (Insecta). Arthropod Syst. Phyl. 77 (1), 125–140.

Polilov, A.A., Ribera, I., Yavorskaya, M.I., Cardoso, A., Grebennikov, V.V., Beutel, R.G., 2019. The phylogeny of Ptiliidae (Coleoptera: Staphylinoidea) – the smallest beetles and their evolutionary transformations. Arthropod Syst. Phyl. 77(3), 433–455.

Qi, X., Lin, X.-L., Ekrem, T., Beutel, R.G., Song, Ch., Orlov, I., Chen, Ch.-T., Wang, X.–H., 2019. A new surface gliding species of Chironomidae: an independent invasion of marine environments and its evolutionary implications. Zool. Scr. 48(1), 81–92.

Richter, A., Keller, R.A., Rosumek, F.B., Economo, E.P., Garcia, F.H., Beutel, R.G., 2019. The cephalic anatomy of workers of the ant species Wasmannia affinis (Formicidae, Hymenoptera, Insecta) and its evolutionary implications. Arthropod Str. Dev. 49, 26–49.

Russell, L.K., Beutel, R.G., 2019. 2.1. Taxonomoy and distribution. In: Beutel, R.G., Friedrich, F. (Eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Nannomecoptera and Neomecoptera. Walter De Gruyter, Berlin, New York.

Tröger, D., Beutel, R.G., Pohl, H., 2019. The abdomen of a free‐living female of Strepsiptera and the evolution of the birth organs. J. Morph. 280(5), 739–755.

Vasilikopoulos, A., Balke, M., Beutel, R.G., Donath, A., Podsiadlowski, L., Pflug, J. M., Waterhourse, R.M., Meusemann, K., Peters, R.S., Escalona, H.E., Mayer, Ch., Liu, Sh., Hendrich, L., Alarie, Y., Bilton, D.T., Jia, F., Zhou, X., Maddison, D.R., Misof, B., 2019. Phylogenomics of the superfamily Dytiscoidea (Coleoptera: Adephaga) with an evaluation of phylogenetic conflict and systematic error. Mol. Phylog. Evol. 135, 270–285.

Yan, E.V., Beutel, R.G., Lawrence, J.F., Yavorskaya, M.I., Hörnschemeyer, T., Pohl, H., Vassilenko, D.V., Bashkuev, A.S., Ponomarenko, A.G., 2019. Archaeomalthus – (Coleoptera, Archostemata) a “ghost adult” of Micromalthidae from Upper Permian deposits of Siberia? Histor. Biol. 32(8), 1019–1027.

Yavorskaya, M.I., Beutel, R.G., Farisenkov, S.E., Polilov, A.A., 2019. The locomotor apparatus of one of the smallest beetles – The thoracic skeletomuscular system of Nephanes titan (Coleoptera, Ptiliidae). Arthropod Str. Dev. 48, 71–82.

Zhou, Y.L., Zhou, H.Zh., Ślipiński, A., Beutel, R.G., 2019. Evolution of a hyper-complex intromittent organ in rove beetles – the endophallus of Xantholinini (Staphylinidae: Coleoptera). Zool. J. Linn. Soc. 188(4), 1277–1295.

Bai, M., Beutel, R.G., Zhang, W., Wang, Sh., Hörnig, M., Gröhn, C. Yan, E., Yang, X.-K., Wipfler, B., 2018. †Caputoraptor elegans – a new Cretaceous insect with a unique cephalo-thoracic scissor device. Curr. Biol.28(3), 438–443.

Batelka, J., Prokop, J. Pohl, H., Bai, M., Zhang, W.W., Beutel, R.G., 2018. Highly specialized Cretaceous beetle parasitoids (Ripiphoridae) identified with optimized visualization of microstructures. Syst. Entomol. 44(2), 396–407.

Beutel, R.G., Pohl, H., Yan, E.V., Anton, E., Liu, S.-P., Ślipiński, S.A., McKenna, D., Friedrich, F., 2018. The phylogeny of Coleopterida (Hexapoda) – morphological characters and molecular phylogenies. Syst. Entomol. 74 (1), 75–102.

Brehm, G., 2018. Revision of the genus Callipia Guenée (1858) (Lepidoptera, Geometridae) with the description of 15 new species. Eur. J. Taxon. 404, 1–54.

Brehm, G., von Knorre, D., Fischer, M.S., 2018. JENA: The Scientific Zoological Collections at the Phyletisches Museum in Jena: Historical Development and Conservational Challenges. In: Zoological Collections of Germany. Springer, pp. 467–476.

Jandausch, K., Beutel, R.G., Pohl, H., Aspöck, U., Winterton, Sh.L., 2018. Morphology of the primary larva of Mantispa aphavexelte (Mantispidae: Neuroptera) and phylogenetic implications to the order of Neuroptera. Arthropod Syst. Phyl. 76(3), 529–560.

Jandausch, K., Beutel, R.G., Pohl, H., Gorb, S.N., Büsse, S., 2018. The legs of “spider associated” parasitic primary larvae of Mantispa aphavexelte (Mantispidae, Neuroptera) –Attachment devices and phylogenetic implications. Arthropod Str. Dev. 47 (5), 449–456.

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Liu, S.P., Friedrich, F., Büsse, S., Petersen, D., Gorb., S., Beutel, R.G., 2018. The thoracic anatomy of the swift lousefly Crataerina pallida (Diptera) – functional implications and character evolution in Hippoboscidea. Zool. J. Linn. Soc. 185 (1), 111–131.

Liu, S.P., Wipfler, B., Beutel, R.G., 2018. The unique locomotor apparatus of whirligig beetles of the tribe Orectochilini (Gyrinidae, Coleoptera). J. Zool. Syst. Evolut. Res. 56 (2), 196–208.

Luo, X.-Zh., Wipfler, B., Ribera, I., Liang, H.-B., Tian M.-Y., Ge, S.-Q., Beutel, R.G., 2018. The thoracic morphology of cave-dwelling and free-living ground beetles from China (Coleoptera, Carabidae, Trechinae). Arthropod Str. Dev. 47(6), 662–674.

Luo, X.-Zh., Wipfler, B., Ribera, I., Liang, H.-B., Tian, M. Y., Ge, S.-Q., Beutel, R.G., 2018. The cephalic morphology of free-living and cave-dwelling species of trechine ground beetles from China (Coleoptera, Carabidae). Org. Divers. Evol. 18(1), 125–142.

Mashimo, Y., Müller, P., Pohl, H., Beutel, R.G., 2018. The “hairy beast” – Zorotypus hirsutus sp.n., an unusual new species of Zoraptera (Insecta) from Burmese amber. Zootaxa 4508(4), 562–568.

Pohl, H., Batelka, J., Prokop, J. Müller, P., Yavorskaya, M., Beutel, R.G., 2018. A needle in a haystack: Mesozoic origin of parasitism in Strepsiptera revealed by first definite Cretaceous primary larva (Insecta). PeerJ 6, e5943https://doi.org/10.7717/peerj.5943External link.

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Yan, E.V., Beutel, R.G., Lawrence, J F., 2018. Whirling in the late Permian: ancestral Gyrinidae show early radiation of beetles before Permian-Triassic mass extinction. BMC Evolut. Biol. 18(1), 33.

Yavorskaya, M.I., Anton, E., Jałoszyński, P., Polilov, A., Beutel, R.G., 2018. Cephalic anatomy of Sphaeriusidae and a morphology‐based phylogeny of the suborder Myxophaga (Coleoptera). Syst. Entomol. 43 (4), 777–797.

Yavorskaya, M.I., Hörnschemeyer, T., Beutel, R.G., 2018. The head morphology of Micromalthus debilis(Coleoptera: Archostemata: Micromalthidae) – an “archaic” beetle with an unusual morphology and a unique life cycle. Arthropod Syst. Phyl. 76(3), 475–486.

Antunes-Carvalho, C., Yavorskaya, M., Gnaspini, P^., Ribera, I., Beutel, R.G., 2017. Cephalic anatomy and three-dimensional reconstruction of the head of Catops ventricosus (Weise, 1877) (Coleoptera: Leiodidae: Cholevinae). Org. Divers. Evol. 17(1), 199–212.

Beck, J., McCain, C.M., Axmacher, J.C., Ashton, L.A., Bärtschi, F., Brehm, G., Choi, S.-W., Cizek, O., Colwell, R.K., Fiedler, K., others, 2017. Elevational species richness gradients in a hyperdiverse insect taxon: a global meta-study on geometrid moths. Glob. Ecol. Biogeogr. 26, 412–424.

Beutel, R.G., Yan, E., Richter, A., Büsse, S., Miller, K.B., Yavorskaya, M., Wipfler, B., 2017. The head of Heterogyrus milloti (Coleoptera: Gyrinidae) and its phylogenetic implications. Arthropod Syst. Phyl. 75(2), 261–280.

Beutel, R.G., Yavorskaya, M., Mashimo, Y., Fukui, M., & Meusemann, K., 2017. The Phylogeny of Hexapoda (Arthropoda) and the Evolution of Megadiversity. Proc. Arthr. Embryol. Soc. Japan 51, 1–15.

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Liu, S.P., Wipfler, B., Niitsu, S., Beutel, R.G., 2017. The thoracic anatomy of the male and female winter moth Nyssiodes lefuarius (Lepidoptera: Geometridae) and evolutionary changes in the thorax of moths and butterflies. Org. Divers. Evol. 17 (3), 565–594.

Neubert, D., Simon, S., Beutel, R.G., Wipfler, B., 2017. The head of the earwig Forficula auricularia (Dermaptera) and its evolutionary implications. Arthropod Syst. Phyl. 75(1), 99–124.

Richter, A., Beutel, R.G., Pohl, H., 2017. The cephalothorax of females of Xenos vesparum (Insecta, Strepsiptera, Xenidae). Arthropod Syst. Phyl. 75(2), 327–347.

Schneeberg, K., Bauernfeind, R., Pohl, H., 2017. Comparison of cleaning methods for delicate insect specimens for scanning electron microscopy. Microsc. Res. Tech. 80, 1199–1204. doi:10.1002/jemt.22917

Strutzenberger, P., Brehm, G., Gottsberger, B., Bodner, F., Seifert, C.L., Fiedler, K., 2017. Diversification rates, host plant shifts and an updated molecular phylogeny of Andean Eois moths (Lepidoptera: Geometridae). PLoS One 12, e0188430.

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Yan, E., Beutel, R.G., Ponomarenko, A.G., 2017. †Ademosynidae (Insecta: Coleoptera) – a new concept for a coleopteran key taxon and its phylogenetic affinities to the extant suborders. Palaeontol. Electr. 20(2), 1–21.

Yan, E., Lawrence, J.F., Beattie, R., Beutel, R.G., 2017. At the dawn of the great rise: †Ponomarenkia belmonthensis (Insecta: Coleoptera), a new remarkable late Permian beetle from the Southern Hemisphere. J. Syst. Palaeontol. 16(7), 611–619.

Yan, E., Ponomarenko, A.G., Beutel, R.G., 2017. Peltosynidae – a new beetle family from the Middle-Late Triassic of Kirghyzia – its affinities to Polyphaga (Insecta, Coleoptera) and the groundplan of the megadiverse suborder. J. Syst. Palaeontol. 16(6), 515–553.

Yavorskaya, M., Beutel, R.G., Polilov, A., 2017. Head morphology of the smallest beetles (Coleoptera: Ptiliidae) and the evolution of sporophagy within Staphyliniformia. Arthropod Syst. Phyl. 75 (3), 417–434.

Zenker, M.M., Wahlberg, N., Brehm, G., Teston, J.A., Przybylowicz, L., Pie, M.R., Freitas, A.V.L., 2017. Systematics and origin of moths in the subfamily Arctiinae (Lepidoptera, Erebidae) in the Neotropical region. Zool. Scr. 46, 348–362.

Anton, E., Yavorskaya, M.I., Beutel, R.G., 2016. The head morphology of adults of Clambidae (Coleoptera: Polyphaga: Scirtoidea). J. Morph. 277, 615–633.

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Bai, M., Beutel, R.G., Klass, K.-D., Zhang, W., Yang, X.-K., Wipfler, B., 2016. †Alienoptera – a new insect order in the roach - mantodean twilight zone. Gondwana Res. 39, 317–326.

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Beutel, R.G., Zhang, W.W., Pohl, H., Wappler, T., Bai, M., 2016. A miniaturized beetle larva in Cretaceous Burmese amber: reinterpretation of a fossil “strepsipteran triungulin”. Ins. Syst. Evol. 47(1), 83–91.

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Dallai, R., Gottardo, M., Beutel, R.G., 2016. Structure and evolution of insect sperm: new interpretations in the age of phylogenomics. Ann. Rev. Entomol. 61, 1–23.

Fabian, B., Schneeberg, K., Beutel, R.G., 2016. Comparative thoracic anatomy of the wild type and wingless (wg1cn1) mutant of Drosophila melanogaster (Diptera). Arthropod Str. Dev. 45(6), 611–636.

Gottardo, M., Dallai, R., Mercati, D., Hörnschemeyer, T., Beutel, R.G., 2016. The evolution of insect sperm – an unusual character system in a megadiverse group. J. Zool. Syst. Evolut. Res. 54(4), 237–256.

Kjer, K.M., Simon, C., Yavorskaya, M., Beutel, R.G., 2016. Progress, pitfalls, and parallel universes: A history of insect phylogenetics. J. R. Soc. Interf. 13, 20160363, http://dx.doi.org/10.1098/rsif.2016.0363.

Knauthe, P., Beutel, R.G., Hörnschemeyer, T., Pohl, H., 2016. Serial block-face scanning electron microscopy sheds new light on the head anatomy of an extremely miniaturized insect larva (Insecta, Strepsiptera). Arthropod Syst. Phyl. 74 (2), 107–126.

Löwe, S., Beutel, R.G., Pohl, H., 2016. The female cephalothorax of Stylops ovinae Noskiewicz & Poluszyński, 1928 (Strepsiptera: Stylopidae). Arthropod Syst. Phyl. 74, 65–81.

Peinert, M., Wipfler, B., Jetschke, G., Kleinteich, T., Gorb, S.N., Beutel, R.G., Pohl, H., 2016. Traumatic insemination and female counter-adaptation in Strepsiptera (Insecta). Sci. Rep. 6, 25052.

Pohl, H., Beutel, R.G., 2016. †Kinzelbachilla ellenbergeri – a new ancestral species, genus and family of Strepsiptera (Insecta). Syst. Entomol. 41 (1): 287–297.

Rodríguez-Castañeda, G., Brehm, G., Fiedler, K., Dyer, L.A., 2016. Ant predation on herbivores through a multitrophic lens: how effects of ants on plant herbivore defense and natural enemies vary along temperature gradients. Curr. Opin. Insect Sci. 14, 73–80.

Wipfler, B., Pohl, H., Yavorskaya, M.I., Beutel, R.G., 2016. A review of methods for analysing insect structures – the role of morphology in the age of phylogenomics. Curr. Opin. Ins. Sci. 18, 60–68.

Bauernfeind, R., Schneeberg, K., Beutel, R.G., 2015. The larval head of Exechia (Mycetophilidae) and Bibio (Bibionidae) (Diptera). Arthropod Str. Dev. 44(4), 326–345.

Bodner, F., Brehm, G., Fiedler, K., 2015. Many caterpillars in a montane rain forest in Ecuador are not classical herbivores. J. Trop. Ecol. 31, 473–476.

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Brehm, G., Fischer, M., Gorb, S., Kleinteich, T., Kühn, B., Neubert, D., Pohl, H., Wipfler, B., Wurdinger, S., 2015. The unique sound production of the Death’s-head hawkmoth (Acherontia atropos (Linnaeus, 1758)) revisited. Sci. Nat. 102, 1–13.

Dallai, R., Gottardo, M., Mercati, D., Rafael, José Albertino, Machida, R., Mashimo, Y., Matsumura, Y. & Beutel, R.G. 2015. The intermediate sperm type and genitalia of Zorotypus shannoni Gurney – evidence supporting infraordinal lineages in Zoraptera (Insecta). Zoomorph. 134(1), 79–91.

Fabian, B., Russell, L., Friedrich, F., & Beutel, R.G., 2015. The morphology of the larval head of the enigmatic boreid Caurinus dectes (Mecoptera). Arthropod Syst. Phyl. 73 (3), 385–399.

Fraulob, M., Beutel, R.G., Machida, R., Pohl, H., 2015. The embryonic development of Stylops ovinae(Strepsiptera, Stylopidae) with emphasis on external morphology. Arthropod Str. Dev. 44 (1), 42–68.

Fraulob, M., Pohl, H., 2015. First record of Eoxenos laboulbenei de Peyerimhoff, 1919 (Strep-siptera, Mengenillidae) from Tuscany, central Italy. Linzer biologische Beiträge 47, 591–595.

Friedemann, K., Gorb, E., Kunert, G., Beutel, R.G., 2015. Attachment forces of pea aphid host races on different legume species. Ecol. Entomol. 40 (6), 732–740.

Ge, S.-Q., Hua, Y., Ren, J., Ślipiński, A., Heming, B., Beutel, R.G., Yang, X.-K., Wipfler, B., 2015. Transformation of head structures during the metamorphosis of Chrysomela populi (Coleoptera, Chrysomelidae). Arthropod Syst. Phyl. 73 (1), 129–152.

Gottardo, M., Vallotto, D., Beutel, R.G., 2015. Giant stick insects reveal unique ontogenetic changes in biological attachment devices. Arthropod Str. Dev. 44 (2), 195–199.

Hädicke, C.W., Blank, S.M., Pohl, H., Müller, C., Sombke, A., 2015. Sensing the world without antennae and eyes: external structure and distribution of sensilla in Eosentomon pinetorum Szeptycki, 1984 and on the protarsus of Acerentomon franzi Nosek, 1965 (Hexapoda: Protura). Soil Organisms 87, 29–49.

Hua, Y., Beutel, R.G., Ge, S.-Q., Yang, X.-K., 2015. The larval head structures of Podagricomela shirahatai (Chûjô) (Chrysomelidae, Galerucinae, Alticini) and morphological effects of leaf mining. J. Morph. 276 (4), 446–457.

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Dallai, R., Gottardo, Mercati, D., Machida, R., Mashimo, Y., Matsumura, Y., Beutel, R.G., 2014. Giant spermatozoa and a huge spermatheca: a case of coevolution of male and female reproductive organs in the ground louse Zorotypus impolitus (Insecta, Zoraptera). Arthropod Str. Dev. 43(2), 135–151.

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Schneeberg, K., Beutel, R.G., 2014. The evolution of head structures in lower Diptera. ScienceOpen, online.

Song, K.Q., Xue, H.J., Beutel, R.G., Bai, M., Bian, D.J., Liu, J., Ruan, Y.Y., Li, W.Z, Jia, F.L. & Yang, X.K. 2014. Habitat-dependent diversification and parallel molecular evolution – water scavenger beetles as a case study. Curr. Zool. 60 (5), 561–570.

Wipfler, B., Bai, M., Schoville, S., Dallai, R., Uchifune, T., Machida, R., Cui, Y., Beutel, R.G., 2014. Ice Crawlers (Grylloblattodea) – the history of the investigation of a highly unusual group of insects. J. Ins. Biodiv. 2(2), 1–25.

Yavorskaya, M.I., Leschen, R.A.B., Polilov, A.A., Beutel, R.G., 2014. Unique rostrate larvae and basidiomycophagy in the beetle family Corylophidae. Arthropod Str. Dev. 43(2), 153–62.

Bai, M., Beutel, R. G., Shih, Ch.-K. Ren, D., & Yang, X.-K., 2013. Septiventeridae, a new and ancestral fossil family of Scarabaeoidea (Insecta: Coleoptera) from the Late Jurassic to Early Cretaceous Yixian Formation. J. Syst. Paleontol. 11(3), 359–374.

Beutel, R.G., Pohl, H., 2013. Insecta (Hexapoda). In: Westheide, W., Rieger, R. (Eds.) Spezielle Zoologie, Teil 1. Einzeller und Wirbellose Tiere. Elsevier Spektrum Akademischer Verlag, München.

Beutel, R.G., Wang, B., Tan, J.-J., Ge, S.-Q., Ren, D., Yang, X.-K., 2013. On the phylogeny and evolution of Mesozoic and extant lineages of Adephaga (Coleoptera, Insecta). Cladistics 29 (2), 147–165.

Beutel, R.G., Wipfler, B., Gottardo, M., & Dallai, R., 2013. Polyneoptera or “lower Neoptera” – new light on old and difficult phylogenetic problems. Atti Accad. Naz. Ital. Entomol. 61, 133–142.

Brehm, G., Strutzenberger, P., Fiedler, K., 2013. Phylogenetic diversity of geometrid moths decreases with elevation in the tropical Andes. Ecography (Cop.). 36, 1247–1253.

Dallai, R., Gottardo, M., Mercati, D., Machida, R., Mashimo, Y., Matsumura, Y., Beutel, R.G., 2013. Divergent mating patterns and a unique mode of external sperm transfer in Zoraptera: an enigmatic group of pterygote insects. Naturwiss. 100, 581–594.

Friedemann, K., Spangenberg, R., Yoshizawa, K., Beutel, R.G., 2013. The evolution of attachment structures in the highly diverse Acercaria (Hexapoda). Cladistics 30(2), 170–201.

Friedrich, F., Beckmann, F., Beutel, R.G., 2013. The head of Merope tuber (Meropeidae) and the phylogeny of Mecoptera (Hexapoda). Arthropod Str. Dev. 42(1), 69–88.

Friedrich, F., Matsumura, Y., Pohl, H., Bai, M., Hörnschemeyer, T., Beutel, R.G., 2013. Insect morphology in the age of phylogenomics – innovative techniques and its future role in systematics. Entomol. Sci. 17 (1), 1–24.

Hörnschemeyer, T., Haug, J., Bethoux, O., Beutel, R.G., Charbonier, S. Thomas A. Hegna, T.A., Koch, M., Rust, S., Wedmann, S., Bradler, S., Willmann, R., 2013. Strudiella not a Devonian insect. Nature 494, E3–E4.

Hua, Y., Ge, S., Beutel, R.G., Zhang, Y., Cui, J., Yang, X.K., 2013. The larvae of Altica koreana (Ogloblin) and A. viridicyanea (Baly) (Coleoptera: Chrysomelidae: Galerucinae: Alticini). Zootaxa, 3694 (5), 461–470.

Hünefeld, F., Brehm, G., Pohl, H., 2013. A simple “hands-off” apparatus to inflate eversible soft parts of the genitalia of small insect specimens. Microsc. Res. Tech. 76, 258–263.

Lawrence, J.F., Ślipiński, A., Beutel, R.G., Newton, A.F., 2013. A possible larva of Lepicerus inaequalis Motschulsky (Coleoptera: Myxophaga: Lepiceridae) from Panama. Zootaxa 3701 (3), 393–400.

Mashimo, Y., Beutel, R.G., Dallai, R., Lee, Ch.-Y, Machida, R., 2013. Embryonic development of Zoraptera with special reference to external morphology, and its phylogenetic implications (Insecta). J. Morph. 275 (3), 295–312.

Mashimo, Y., Yoshizawa, K., Engel, M.S., Ghani, A.B., Dallai, R., Beutel, R.G., Machida, R. 2013. Zorotypus in Peninsular Malaysia (Zoraptera: Zorotypidae), with the description of three new species. Zootaxa 3717 (4), 498–514.

Matsumura, Y., Machida, R., Wipfler, B., Beutel, R.G., & Yoshizawa, K., 2013. Parallel evolution of novelties: extremely long intromittent organs in the leaf beetle subfamily Criocerinae. Evol. Dev. 13(4), 305–315.

Onmaz, A.C., Beutel, R., Schneeberg, K., Pavalaoiu, A.N., Komarek, A., v.d. Hoven, R., 2013. Vectors and vector-borne diseases of horses. Vet. Res. Comm. 37(1), 65–81.

Pohl, H. & Beutel, R.G., 2013. The Strepsiptera-Odyssey: the history of the systematic placement of an enigmatic parasitic insect order. – Entomologia 1: 17–28.

Pohl, H., Dallai, R., Gottardo, M., Beutel, R.G., 2013. The spermatozoon of Mengenilla moldrzyki (Strepsiptera, Mengenillidae): ultrastructure and phylogenetic considerations. Arthropod Str. Dev. 45, 446– 451.

Russell, L.K., Dallai, R., Gottardo^, M., Beutel, R.G., 2013. The sperm ultrastructure of Caurinus dectes Russell (Mecoptera: Boreidae) and its phylogenetic implications. Tiss. Cell 45 (6), 397–401.

Schneeberg, K. Harris, M., Polilov, A., & Beutel, R.G. 2013. The adult head morphology of the Hessian fly Mayetiola destructor (Diptera, Cecidomyiidae). J. Morph. 274 (11), 1299–1311.

Schneeberg, K., Krause, K., Beutel, R.G., 2013. The adult head of Axymyia furcata (Axymyiidae, Diptera). Arthropod Syst. Phyl. 71(2), 91–102.

Spangenberg, R., Friedemann, K., Weirauch, C., Beutel, R.G., 2013 The head morphology of the potentially basal heteropteran lineages Enicocephalomorpha, Dipsocoromorpha and Gerromorpha (Hemiptera). Arthropod Syst. Phyl. 71 (2), 103–136.

Spangenberg, R., Wipfler, B., Friedemann, K., Pohl, H., Weirauch, C., Hartung, V., Beutel, R.G., 2013. The cephalic morphology of the Gondwanan key taxon Hackeriella (Coleorrhyncha, Hemiptera). Arthropod Str. Dev. 42, 315–337.

Werner, F.A., Jantz, N., Krashevska, V., Peters, T., Behling, H., Maraun, M., Scheu, S., Brehm, G., 2013. Climate change: effects on biodiversity and ecosystem functioning. In: Ecosystem Services, Biodiversity and Environmental Change in a Tropical Mountain Ecosystem of South Ecuador. Springer, pp. 247–263.

Wipfler, B., Schneeberg, K., Löffler, A., Hünefeld, F., Meier, R., Beutel, R.G., 2013. The skeletomuscular system of the larva of Drosophila melanogaster (Drosophilidae, Diptera) – a contribution to the morphology of a model organism. Arthropod Str. Dev. 42(1), 47–68.

Anton, E., Beutel, R.G., 2012. The head morphology of Dascillus (L.) (Dascilloidea: Dascillidae) and GlaresisErichson (Scarabeoidea: Glaresidae) and its phylogenetic implications. Arthropod Syst. Phyl.70(1), 3–42.

Bai, M., Beutel, R.G., Song, K.-Q., Liu, W.-G., Malqin, H., Li, Sh., Hu, X.-Y., Yang, X.-K., 2012 Evolutionary patterns of hind wing morphology in dung beetles (Coleoptera: Scarabaeinae). Arthropod Str. Dev.41(5), 505–513.

Beutel, R.G., Kristensen, N.P., 2012. Morphology and insect systematics in the era of phylogenomics. Arthropod Str. Dev.41, 303–305.

Blanke, A., Greve, C., Wipfler, B., Beutel, R., Holland, B., Misof, B., 2012. The identification of concerted convergence in insect heads corroborates Palaeoptera. Syst. Biol. 162 (2), 250–263.

Blanke, A., Wipfler, B., Letsch, H., Koch, M., Beutel, R., Misof, B., 2012. Revival of Palaeoptera – head characters support a monophyletic origin of Odonata and Ephemeroptera (Insecta). Cladistics 28(6), 560–581.

Bodner, F., Strutzenberger, P., Brehm, G., Fiedler, K.2012. Species richness and host specificity among caterpillar ensembles on shrubs in the Andes of southern Ecuador. Neotrop. Entomol. Publ. online.

Dallai, R., Mercati, D., Gottardo, M, Dossey, A.T., Machida, R., Mashimo, Y., Beutel, R.G., 2012. The male and female reproductive systems of Zorotypus hubbardi Caudell, 1918 (Zoraptera). Arthropod Str. Dev.41(4), 337–359.

Dallai, R., Mercati, D., Gottardo, M., Machida, R., Mashimo, Y., Beutel, R.G., 2012. The fine structure of the female reproductive system of Zorotypus caudelli Karny (Zoraptera). Arthropod Str. Dev.41, 51–63.

Fraulob, M. Wipfler, B., Hünefeld, F., Pohl, H., Beutel, R.G., 2012. The larval abdomen of the enigmatic Nannochoristidae (Mecoptera, Insecta). Arthropod Str. Dev. 41 (2), 187–198.

Ge, S.-Q., Daccordi, M., Beutel, R.G., Zhi, C.-J., Li, W-ZH., Yang, X.-K., 2012. Revision of the Eastern Asian genera Ambrostoma Motschulsky and Parambrostoma Chen (Coleoptera: Chrysomelidae: Chrysomelinae). Syst. Entomol.37, 332–345.

Ge, S.-Q., Wipfler, B., Pohl, H., Hua, Y., Ślipiński, A., Yang, X.-K., Beutel, R.G., 2012. The first complete 3D reconstruction of a Spanish Fly primary larva (Lytta vesicatoria, Meloidae, Coleoptera). PLOS ONE 7(12), e52511.

Hünefeld, F., Beutel, R.G., 2012. The female postabdomen of the enigmatic Nannochoristidae (Insecta: Mecopterida) and its phylogenetic significance. Acta Zool. 93, 231–238.

Hünefeld, F., Friedrich, F., Hörnschemeyer, T., Pohl, H., Beutel, R.G., 2012. The phylogeny of Holometabola and the role of morphology in the age of phylogenomics. DGaaE Nachrichten Vol. 18 (2012).

Hünefeld, F., Mißbach, C., Beutel, R.G., 2012. The morphology and evolution of the female postabdomen of Holometabola (Insecta). Arthropod Str. Dev.41(4), 361–371.

Jałoszyński, P., Beutel, R.G., 2012 Functional morphology and evolution of specialized mouthparts of Cephenniini (Scydmaeninae, Staphylinidae). Arthropod Str. Dev. 41(6), 593–607.

Koeth, M., Friedrich, F., Pohl, H., Beutel, R.G., 2012. The thoracic skeleto-muscular system of Mengenilla(Strepsiptera: Mengenillidae) and its phylogenetic implications. Arthropod Str. Dev. 41(4), 323–335.

Kukalová-Peck, J., Beutel, R.G., 2012. Is the Carboniferous †Adiphlebia lacoana really the “oldest beetle”? Critical reassessment and description of a new Permian beetle family. Eur. J. Entomol. 109(4), 633–645.

Letsch, H.O., Meusemann, K., Wipfler, B., Schütte, K., Beutel, R., Misof, B., 2012. Insect phylogenomics: results, problems and the impact of matrix composition. Proc. R. Soc. B 279 (1741), 3282–3290.

Niehuis, O., Hartig, G., Grath, S., Pohl, H., Lehmann, J., Tafer, H., Donath, A., Krauss, V., Eisenhardt, C., Hertel, J., Petersen, M., Mayer, C., Meusemann, K., Peters, R.S., Stadler, P.F., Beutel, R.G., Bornberg-Bauer, E., McKenna, D.D., Misof, B., 2012. Genomic and morphological evidence converge to resolve the enigma of Strepsiptera. Curr. Biol. 22 (14), 1309–1313.

Pohl, H., Niehuis, O., Gloyna, K., Misof, B., Beutel, R.G., 2012. A new species of Mengenilla (Insecta, Strepsiptera) from Tunisia. ZooKeys 198, 79.

Ribera, I., Beutel, R.G., 2012. Capítulo 28. Héxapodos. Pp. 280–291 In: Vargas, P., Zardoya, R. (Eds.), El Árbol de la Vida: Systemática y evolución de los seres vivos. Impulso Global Solutions, Madrid.

Ribera, I., Beutel, R.G., 2012. Capítulo 31. Coleópteros. Pp. 312–321 In: Vargas, P., Zardoya, R. (Eds.), El Árbol de la Vida: Systemática y evolución de los seres vivos. Impuslo Global Solutions, Madrid.

Schneeberg, K., Friedrich, F., Courtney, G.W., Wipfler, B., Beutel, R.G., 2012. The larvae of Nymphomyiidae (Diptera, Insecta) – ancestral and highly derived? Arthropod Str. Dev.41(3), 293–301.

Strutzenberger, P., Brehm, G., Fiedler, K., 2012. DNA barcode sequencing from old type specimens as a tool in taxonomy: a case study in the diverse genus Eois (Lepidoptera: Geometridae). PLoS One 7, e49710.

Trautwein, M.D., Wiegmann, B.M., Beutel, R., Kjer, K., Yeates, D.K., 2012. Advances in insect phylogeny at the dawn of the postgenomic era. An. Rev. Entomol. 57, 449–68.

Wipfler, B., Courtney, G.W., Craig, D.A, Beutel, R.G., 2012. First µ-CT based 3D reconstruction of a dipteran larva – the head morphology of Protanyderus (Tanyderidae) and its phylogenetic implications. J. Morph. 273(9), 968-980.

Wipfler, B., Pohl, H., Predel, R., 2012. Two new genera and two new species of Mantophasmatodea (Insecta, Polyneoptera) from Namibia. – ZooKeys 166: 75–98.

Aguirre, L.F., Anderson, E.P., Brehm, G., Herzog, S.K., Joergensen, P.M., Kattan, G.H., Maldonado, M., Martínez, R., Mena, J.L., Pabón, J.D., Seimon, A., Toledo, C., 2011. Phenology and interspecific Ecological Interactions of Andean Biota in the Face of Climate Change. In: Herzog, S.K., Martínez, R., Joergensen, P.M., Tiessen, H. (Eds.), Climate Change and Biodiversity in the Tropical Andes. Inter-American Institute of Global Change Research and Scientific Committee on Problems of the Environment. Sao José dos Campos, Brazil, and Paris, France, pp. 68–92.

Anton, E., Niederegger, S., Beutel, R.G., 2011. Beetles and flies collected on pig carrion in an experimental setting in Thuringia and their forensic implications. Med. Vet. Entomol. 25(4), 353-364.

Beck, J., Brehm, G., Fiedler, K., 2011. Links between the environment, abundance and diversity of Andean moths. Biotropica 43, 208–217.

Beutel, R.G., Friedrich, F., Hörnschemeyer, T., Pohl, H., Hünefeld, F., Beckmann, F., Meier, R., Misof, B., Whiting, M.F., Vilhelmsen, L., 2011. Morphological and molecular evidence converging upon a robust phylogeny of the megadiverse Holometabola. Cladistics 26, 1–15.

Brehm, G., 2011. 19: Diversity of geometrid moths in two Neotropical rain forests. Trop. Mont. Cloud For. Sci. Conserv. Manag. 192.

Brehm, G., Bodner, F., Strutzenberger, P., Hünefeld, F., Fiedler, K., 2011. Neotropical Eois (Lepidoptera: Geometridae): checklist, biogeography, diversity, and description patterns. Ann. Entomol. Soc. Am. 104, 1091–1107.

Dallai, R., Mercati, D., Gottardo, M., Machida, R., Mashimo, Y., Beutel, R.G., 2011. The male reproductive system of Zorotypus caudelli Karny (Zoraptera): sperm structure and spermiogenesis. Arthropod Str. Dev.40 (6), 531-47.

Dressler, C., Ge, S.-Q., Beutel, R.G., 2011. Is Meru a specialised noterid (Coleoptera, Adephaga)? Syst. Entomol.36 (4), 705–712.

Friedemann, K. Wipfler, B., Bradler, S., Beutel, R. G., 2012. On the head morphology of Phyllium and the phylogenetic relationships of Phasmatodea (Insecta). Acta Zool. 93(2), 184–199.

Ge, S.-Q., Daccordi, M, Beutel, R.G., Li W.-Z., Yang, X.-K., 2011. Revision of the chrysomeline genera Potaninia, Suinzona and Taipinus (Coleoptera) from eastern Asia, with a biogeographic scenario for the Hengduan mountain region in south-western China. Syst. Entomol.36 (4), 644–671.

Ge, S.-Q., Hörnschemeyer, T., Friedrich, F., Beutel, R.G., 2011. Is Crowsoniella relicta really a cucujiform beetle? Syst. Entomol.36(1), 175–179.

Hünefeld, F., Marvaldi, A., Müller, B., Lawrence, J.F., Beutel, R.G., 2011. The male postabdomen of the “ancestral” archostematan beetle Tetraphalerus bruchi Heller, 1913 (Ommatidae) and its phylogenetic significance. Arthropod Str. Dev.40 (23), 146–158.

Hünefeld, F., Pohl, H., Wipfler, B., Beckmann, F., Beutel, R.G., 2011. The male postabdomen and genital apparatus of †Mengea tertiaria, a strepsipteran amber fossil (Insecta). J. Zool. Syst. Evolut. Res. 49(4), 298–308.

Jałoszyński, P., Hünefeld, F., & Beutel, R.G., 2011. The evolution of “deformed” brains in ant-like stone beetles (Scydmaeninae, Staphylinidae). Arthropod Str. Dev.41(1):17–28.

Larsen, T.H., Brehm, G., Navarrete, H., Franco, P., Gomez, H., Mena, J.L., Morales, V., Argollo, J., Blacutt, L., Canhos, V., 2011. Range shifts and extinctions driven by climate change in the tropical Andes: synthesis and directions. In: Herzog, S.K., Martínez, R., Joergensen, P.M., Tiessen, H. (Eds.), Climate Change and Biodiversity in the Tropical Andes. Inter-American Institute of Global Change Research and Scientific Committee on Problems of the Environment. Sao José dos Campos, Brazil, and Paris, France, pp. 47–67.

Laurance, W.F., Useche, D.C., Shoo, L.P., Herzog, S.K., Kessler, M., Escobar, F., Brehm, G., Axmacher, J.C., Chen, I.-C., Gámez, L.A., others, 2011. Global warming, elevational ranges and the vulnerability of tropical biota. Biol. Conserv. 144, 548–557.

Mashimo, Y., Machida, R., Dallai, R. Gottardo, M., Beutel, R.G., 2011. Egg structure of Zorotypus caudelli Karny (Insecta, Zoraptera, Zorotypidae). Tiss. Cell 43 (4), 230–237.

Schneeberg, K., Courtney, G.W., Beutel, R.G., 2011. Adult head structures of Deuterophlebiidae (Insecta), a highly derived “ancestral” dipteran lineage. Arthropod Str. Dev.40(1), 93–104.

Sihvonen, P., Mutanen, M., Kaila, L., Brehm, G., Hausmann, A., Staude, H.S., 2011. Comprehensive molecular sampling yields a robust phylogeny for geometrid moths (Lepidoptera: Geometridae). PLoS One 6, e20356.

Spangenberg, R., Hünefeld, F., Schneeberg, K., Beutel, R.G., 2012. The male postabdomen and reproductive system of Bibio marci Linnaeus, 1758 (Hexapoda: Diptera: Bibionidae). J. Zool. Syst. Evolut. Res. 50(4), 264–288.

Strutzenberger, P., Brehm, G., Fiedler, K., 2011. DNA barcoding based species delimitation increases species richness count of (Geometridae) moths in a well-studied tropical mountain forest by up to 50%. Insect Sci. 18, 349–62.

Wipfler, B., Machida, R., Müller, B., Beutel, R.G., 2011. On the head morphology of Grylloblattodea (Insecta) and the systematic position of the order – with a new nomenclature for the head muscles of Neoptera. Syst. Entomol.36, 241–266.

Zarinkamar, N., Yang, X., Bao, R., Friedrich, F., Beutel, R., Friedrich, M., 2011. The Pax gene eyegone facilitates repression of eye development in Tribolium. EvoDevo. 2 (1), 8.

Beutel, R.G., Balke, M., Ribera, I. 2010. 3.1. Aspidytidae Ribera, Beutel, Balke & Vogler, 2002. In: Leschen, R.A.B, Beutel, R.G., Lawrence, J.F. (Vol. Eds.), Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 39. Coleoptera, Vol. 2: Morphology and Systematics (Elateroidea, Cucujifomia excl. Phytophaga). Walter De Gruyter, Berlin, New York.

Beutel, R.G., Friedrich, F., Aspöck, U., 2010. The larval head of Nevrorthidae and the phylogeny of Neuroptera (Insecta). Zool. J. Linn. Soc. 158, 533–562.

Beutel, R.G., Schneeberg, K., 2010. Beetles in Darwin’s life and work. Ann. Hist. and Philos. Biol. 15, 142–157.

Beutel, R.G., Zimmermann, D., Krauß, M., Randolf, S., Wipfler, B., 2010. Head morphology of Osmylus fulvicephalus (Osmylidae, Neuroptera) and its phylogenetic implications. Org. Divers. Evol. 10, 311–329.

Bodner, F., Brehm, G., Homeier, J., Strutzenberger, P., Fiedler, K., 2010. Caterpillars and host plant records for 59 species of Geometridae (Lepidoptera) from a montane rainforest in southern Ecuador. J. insect Sci. 10, 67.

Dressler, C., Beutel, R.G., 2010. The morphology and evolution of the adult head of Adephaga (Insecta, Coleoptera). Arthropod Syst. Phyl. 68, 239–287.

Friedrich, F., Beutel, R.G., 2010a. The thoracic morphology of Nannochorista (Nannochoristidae) and its implications for the phylogeny of Mecoptera and Antliophora. J. Zool. Syst. Evol. Res. 48, 50–74.

Friedrich, F., Beutel, R.G., 2010b. Good bye Halteria? The thoracic morphology of Endopterygota (Insecta) and its phylogenetic implications. Cladistics 26, 579–612.

Ge, S.-Q., Friedrich, F., Beutel, R.G., 2010. On the systematic position and taxonomic rank of the extinct myxophagan †Haplochelus (Coleoptera). Insect Systematics and Evolution 41 (4), 329–338.

Lawrence, J. F., Beutel, R.G., Leschen, R.A.B., Ślipiński, S.A., 2010. 2. Glossary of morphological terms. In: Leschen, R.A.B, Beutel, R.G., Lawrence, J.F. (Vol. Eds.), Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 39. Coleoptera, Vol. 2: Morphology and Systematics (Elateroidea, Cucujifomia excl. Phytophaga). Walter De Gruyter, Berlin, New York.

Leschen, R.A.B., Beutel, R.G., 2010. 14. Elateriformia. Introduction, Phylogeny. In Beutel, R.G.& Leschen, R.A.B (Vol. Eds.), Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Leschen, R.A.B., Beutel, R.G., Lawrence, J.F. (vol. eds), 2010. Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 39. Coleoptera, Vol. 2: Morphology and Systematics (Elateroidea, Cucujifomia excl. Phytophaga). Walter De Gruyter, Berlin, New York.

Longhorn, S., Pohl, H. & Vogler, A., 2010. Ribosomal protein genes of holometabolan insects reject the Halteria, instead revealing a close affinity of Strepsiptera with Coleoptera. – Mol. Phyl. Evol. 55, 846–859.

Pohl, H., 2010. A scanning electron microscopy specimen holder for viewing different an­gles of a single specimen. – Micr. Res. Tech. 73, 1073–1076.

Pohl, H., Wipfler, B., Grimaldi, D., Beckmann, F., Beutel, R.G., 2010. Reconstructing the anatomy of the 42 million-year-old fossil †Mengea tertiara (Insecta, Strepsiptera). Naturwiss, 97, 855–859.

Polilov, A.A., Beutel, R.G., 2010. Developmental stages of the hooded beetle Sericoderus lateralis (Coleoptera: Corylophidae) with comments on the phylogenetic position and effects of miniaturization. Arthropod Str. Dev. 39, 52–69.

Rodríguez-Castañeda, G., Dyer, L.A., Brehm, G., Connahs, H., Forkner, R.E., Walla, T.R., 2010. Tropical forests are not flat: How mountains affect herbivore diversity. Ecol. Lett. 13, 1348–1357.

Schneeberg, K., Beutel, R.G., 2010. The adult head structures of Tipulomorpha (Diptera, Insecta) and their phylogenetic implications. Acta Zool. 92 (4), 316–343.

Seago, A., Beutel, R.G., 2010. 11.15. Prostomidae G.G. Thomson, 1859 Spangler & Steiner 2005. In: Leschen, R.A.B, Beutel, R.G., Lawrence, J. F. (Vol. Eds.), Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 39. Coleoptera, Vol. 2: Morphology and Systematics (Elateroidea, Cucujifomia excl. Phytophaga). Walter De Gruyter, Berlin, New York.

Strutzenberger, P., Bodner, F., Brehm, G., Fiedler, K., 2010. Molecular phylogeny of Eois: historical signal of wing patterns and host plant use in a group of species rich tropical moths. Zool. Scr. 39, 609–620.

Axmacher, J.C., Brehm, G., Hemp, A., Tünte, H., Lyaruu, H.V.M., Müller-Hohenstein, K., Fiedler, K., 2009. Determinants of diversity in afrotropical herbivorous insects (Lepidoptera: Geometridae): plant diversity, vegetation structure or abiotic factors? J. Biogeogr. 36, 337–349.

Bernard, D., Ribera, I, Komarek, A., Beutel, R.B., 2009. Phylogenetic analysis of Hydrophiloidea (Coleoptera, Polyphaga) based on molecular data and morphological characters of adults and immature stages. Ins. Syst. Evol. 40, 3–41.

Beutel, R.G., Kristensen, N.-P., Pohl, H., 2009. Resolving insect phylogeny: The significance of cephalic structures of the Nannomecoptera in understanding endopterygote relationships. Arthropod Str. Dev. 38 (5), 427–460.

Beutel, R.G., Leschen, R.A.B., Friedrich, F., 2009. Darwin, Beetles and Phylogenetics. Naturwiss. 96, 1293–1312.

Bravo, F., Pohl, H., Neto, A., Beutel, R.G., 2009. Bahiaxenidae, a “living fossil” and a new family of Strepsiptera (Hexapoda) discovered in Brazil. Cladistics 25, 614–623.

Eberhard, M.J.B, Pass, G., Picker, M.D., Beutel, R., Predel, R., Gorb, S.N., 2009. Structure and function of the arolium of Mantophasmatodea (Insecta). J. Morph. 270 (10), 1247–1261.

Friedrich, F., Farrell, B.D., Beutel, R.G., 2009. The thoracic morphology of Archostemata and the relationships of the extant suborders of Coleoptera (Hexapoda). Cladistics 25, 1–37.

Neugart, C., Schneeberg, K., Beutel, R.G., 2009. The morphology of the larval head of Tipulidae (Diptera, Insecta) – the dipteran groundplan and evolutionary trends. Zool. Anz. 248, 215–235.

Osswald, J., Pohl, H., Beutel, R.G., 2009. Extremely miniaturised and highly complex: The thoracic morphology of the first instar larva of Mengenilla chobauti (Insecta, Strepsiptera). Arthropod Str. Dev. 39 (4), 287–304.

Pohl, H., 2009. The oldest fossil strepsipteran larva (Insecta: Strepsiptera) from the Geisel Valley, Germany (Eocene). Ins. Syst. Evol. 40: 333–347.

Polilov, A.A., Beutel, R.G., 2009. Miniaturization effects in developmental stages of Mikado sp. (Coleoptera: Ptiliidae), one of the smallest free-living insects. Arthropod Str. Dev. 38, 247–270.

Xiaoyun, Y., Weber, M., Zarin-Kamar, N., Posnien, N., Friedrich, F., Wiegand, B., Beutel, R., Damen, W., Bucher, G., Klingler, M., Friedrich, M., 2009. Probing the Drosophila retinal determination gene network in Tribolium (II): the Pax6 genes eyeless and twin of eyeless. Dev. Biol. 333 (1), 215–27.

Balke, M., Ribera, I., Beutel, R., Viloria, A., Garcia, M., Vogler, A.P., 2008. Systematic placement of the recently discovered beetle family Meruidae (Coleoptera: Dytiscoidea) based on molecular data. Zool. Scr. 37 (6),647–650.

Beutel, R.G., Baum, E., 2008. A longstanding entomological problem finally solved? Head morphology ofNannochorista (Mecoptera, Insecta) and possible phylogenetic implications. J. Zool. Syst. Evolut. Res.46 (4), 346–367.

Beutel, R.G., Friedrich, F., 2008. Comparative study of larval head structures of Megaloptera (Hexapoda). Eur. J. Entomol. 105, 917–938.

Beutel, R.G., Friedrich, F., 2008. The phylogeny of Archostemata (Coleoptera) and new approaches in insect morphology. Entomol. Gen. 31 (2), 141–154.

Beutel, R.G., Friedrich, F., Whiting, M.F., 2008. Head morphology of Caurinus (Boreidae, Mecoptera) and its phylogenetic implications. Arthropod Str. Dev. 37, 418–433.

Beutel, R.G., Ge, S.-Q., Hörnschemeyer, T., 2008. On the head morphology Tetraphalerus, the phylogeny of Archostemata and the basal branching events in Coleoptera. Cladistics 23, 1–29.

Beutel, R.G., Ge, S.-Q., Yang, X.Q., 2008. The larval head of Raphidia (Raphidioptera, Insecta) and its phylogenetic significance. Zoology 11, 89–113.

Beutel, R.G., Gorb, S., 2008. Evolutionary scenarios for unusual attachment devices of Phasmatodea and Mantophasmatodea (Insecta). Syst. Entomol. 33, 501–510.

Beutel, R.G., Krogmann, L., Vilhelmsen, L.T., 2008. The larval head morphology of Xyela sp. (Xyelidae, Hymenoptera) and its phylogenetic implications. J. Zool. Syst. Evolut. Res. 46(2), 118–132.

Beutel, R.G., Ribera, I., Bininda-Emonds, O., 2008. A genus-level supertree of Adephaga (Coleoptera). Organisms, Diversity and Evolution 7, 255–269.

Brehm, G., 2008. 15 Patterns of Arctiid Diversity. In Conner WE (ed) Tiger Moths and Woolly Bears: Behavior, Ecology, and Evolution of the Arctiidae, pp. 223–232. Oxford University Press.

Brehm, G., Homeier, J., Fiedler, K., Kottke, I., Illig, J., Nöske, N.M., Werner, F.A., Breckle, S.W., 2008. Mountain rain forests in southern Ecuador as a hotspot of biodiversity--limited knowledge and diverging patterns. In: Gradients in a Tropical Mountain Ecosystem of Ecuador. Springer, pp. 15–23.

Colwell, R.K., Brehm, G., Cardelús, C.L., Gilman, A.C., Longino, J.T., 2008. Global warming, elevational range shifts, and lowland biotic attrition in the wet tropics. Science 322, 258–261.

Fiedler, K., Brehm, G., Hilt, N., Sussenbach, D., Hauser, C.L., 2008. Variation of diversity patterns across moth families along a tropical altitudinal gradient. Ecol. Stud. 198, 167.

Fiedler, K., Brehm, G., Hilt, N., Süssenbach, D., Onore, G., Bartsch, D., Racheli, L., Häuser, C.L., others, 2008. Lepidoptera: Arctiidae, Geometridae, Hedylidae, Pyraloidea, Sphingidae, Saturniidae, and Uraniidae (moths) – checklist Reserva Biológica San Francisco (Prov. Zamora-Chinchipe, S. Ecuador). In: Gradients in a Tropical Mountain Ecosystem of Ecuador. Springer, pp. 155–214.

Friedrich, F., Beutel, R.G., 2008. The thorax of Zorotypus (Hexapoda, Zoraptera) and a new nomenclature for the musculature of Neoptera. Arthropod Str. Dev. 37, 29–54.

Krogmann, L., v. Knorre, D., Beutel, R.G., 2008. Die Chalcidoidea-Sammlung von Ferdinand Rudow (1840–1920) im Phyletischen Museum (Jena). Mitt. Hamburger zool. Mus. Inst. 104, 129–140.

Nöske, N.M., Hilt, N., Werner, F.A., Brehm, G., Fiedler, K., Sipman, H.J.M., Gradstein, S.R., 2008. Disturbance effects on diversity of epiphytes and moths in a montane forest in Ecuador. Basic Appl. Ecol. 9, 4–12.

Pohl, H., Beutel, R.G., 2008. The evolution of Strepsiptera. Zoology 111, 318–338.

Vilhelmsen, L., Turrisi, G.F., Beutel, R.G., 2008. Distal leg morphology, subgenual organs and host detection in Stephanidae (Insecta, Hymenoptera). J. Nat. Hist. 42, 1649–1663.

Baum, E., Beutel, R.G., Hertel, W., 2007. The head capsule, central nervous system and head circulatory system of an aberrant orthopteran, Prosarthria teretrirostris (Caelifera, Hexapoda). Zoology 110, 147–160.

Baum, E., Dressler, C., Beutel, R.G., 2007. Head structures of Karoophasma sp. (Mantophasmatodea) with phylogenetic implications. J. Zool. Syst. Evolut. Res. 45, 104–119.

Beutel, R.G., Bocak, L., Bocakova, M., 2007. Are Polyphaga (Coleoptera) really a basal neopteran lineage – a reply to Kazantsev. Acta Zool. 88, 153–158.

Beutel, R.G., Vilhelmsen, L.B., 2007. Head anatomy of Xyelidae (Hexapoda: Hymenoptera) and phylogenetic implications. Org. Divers. Evol. 7, 207–230.

Brehm, G., 2007. Contrasting patterns of vertical stratification in two moth families in a Costa Rican lowland rain forest. Basic Appl. Ecol. 8, 44–54.

Brehm, G., Colwell, R.K., Kluge, J., 2007. The role of environment and mid-domain effect on moth species richness along a tropical elevational gradient. Glob. Ecol. Biogeogr. 16, 205–219.

Brehm, G., Hartmann, T., Willmott, K.R., 2007. Pyrrolizidine alkaloids and pharmacophagous Lepidoptera visitors of Prestonia amabilis (Apocynaceae) in a montane rain forest in Ecuador. Ann. Missouri Bot. Gard. 94, 465–475.

Fiedler, K., Hilt, N., Brehm, G., Schulze, C.H., 2007. Moths at tropical forest margins – how mega-diverse insect assemblages respond to forest disturbance and recovery. In: Stability of Tropical Rainforest Margins. Springer, pp. 37–58.

Fontaine, B., Bouchet, P., Van Achterberg, K., Alonso-Zarazaga, M.A., Araujo, R., Asche, M., As­poeck, U., Audisio, P., Aukema, B., Bailly, N., Balsamo, M., Bank, R.A, Barnard, P., Belfiore, C., Bogdanowicz, W., Bongers, T., Boxshall, G., Burckhardt, D., Camicas, J.-L., Chylarecki, P., Crucitti, P., Deharveng, L., Dubois, A., Enghoff, H., Faubel, A., Fo­chetti, R., Gargominy, O., Gibson, D., Gib­son, R., Gomez Lopez, M.S., Goujet, D., Harvey, M.S., Heller, K.-G., Van Helsdingen, P., Hoch, H., De Jong, H., De Jong, Y., Karsholt, O., Los, W., Lundquist, L., Magowski, W., Manconi, R., Mar­tens, J., Massard, J.A., Massard-Geimer, G., Mcinnes, S.J., Mendes, L..F, Mey, Eberhard, Michel­sen, Verner, Minelli, Alessandro, Nielsen, Claus, Nafria, Juan M. Nieto, Van Nieukerken, E.J., Noyes, J., Pape, T., Pohl, H., De Prins, W., Ramos, M., Ricci, C., Roselaar, C., Rota, E., Schmidt-Rhaesa, A., Segers, H., Strassen, R. zur, Szeptycki, A., Thibaud, J.-M., Thomas, A., Timm, T., Van Tol, J., Ver­voort, W. & Willmann, R., 2007. The Euro­pean Union's 2010 target: Putting rare spe­cies in fo­cus. Biological Conservation 139: 167–185.

Ge, S.-Q., Beutel, R.G., Yang, X.-K., 2007. Thoracic morphology of adults of Derodontidae and Nosodendridae and its phylogenetic implications (Coleoptera). Syst. Entomol. 32, 635–667.

Häuser, C.L., Fiedler, K., Bartsch, D., Brehm, G., Kling, M., Süssenbach, D., Onore, G., 2007. Butterflies (Lepidoptera: Papilionoidea) – Checklist Reserva Biológica San Francisco (Prov. Zamora-Chinchipe, S. Ecuador). Ecotropical Monogr. 4, 145–154.

Hilt, N., Brehm, G., Fiedler, K., 2007. Temporal Dynamics of Rich Moth Ensembles in the Montane Forest Zone in Southern Ecuador. Biotropica 39, 94–104.

Knorre, D. v., Beutel, R., 2007. Die geowissenschaftlichen Sammlungen im Phyletischen Museum in Jena. Beitr. Geol. Thüringen, NF 13, 61–72.

Komarek, A., Beutel, R.G., 2007. Phylogenetic analysis of Anacaenini (Coleoptera: Hydrophilidae: Hydrophilinae) based on morphological characters of adults. Syst. Entomol. 32, 205–226.

Komarek, A., Beutel, R.G., 2007. Problems in taxonomy and suggestions for a standardized description of new insect taxa. Entomol. Probl. 36 (2), 55–70.

Anton, E., Beutel, R.G., 2006. On the head morphology of Lepiceridae (Coleoptera: Myxophaga) and the systematic position of the family and suborder. Eur. J. Entomol. 103, 85–95.

Bernhard, D., Schmidt, C., Korte, A., Fritzsch, G., Beutel, R.G., 2006. From terrestrial to aquatic habitats and back again – molecular insights into the evolution and phylogeny of Hydrophiloidea (Coleoptera) using multigene analyses. Zool. Scr. 35, (6) 597–306.

Beutel, R.G., Balke, M., Steiner, W.E., 2006. On the systematic position of Meruidae (Coleoptera, Adephaga) and the phylogeny of the smaller hydradephagan families. Cladistics 22, 102–131.

Beutel, R.G., Gorb, S., 2006. A revised interpretation of the evolution of attachment structures in Hexapoda (Arthropoda), with special emphasis on Mantophasmatodea. Arthropod Syst. Phyl. 64 (1), 3–25.

Beutel, R.G., Pohl, H., 2006. Endopterygote systematics – where do we stand and what is the goal (Hexapoda, Arthropoda). Syst. Entomol.31, 202–219.

Brehm, G., Axmacher, J.C., 2006. A comparison of manual and automatic moth sampling methods (Lepidoptera: Arctiidae, Geometridae) in a rain forest in Costa Rica. Environ. Entomol. 35, 757–764.

Friedrich, F., Beutel, R.G., 2006. The pterothoracic skeletomuscular system of Scirtoidea (Coleoptera: Polyphaga) and its implications for the relationships of the beetle suborders. J. Zool. Syst. Evolut. Res. 44 (4), 290–315.

Hilt, N., Brehm, G., Fiedler, K., 2006. Diversity and ensemble composition of geometrid moths along a successional gradient in the Ecuadorian Andes. J. Trop. Ecol. 22, 155–166.

Arce-Pérez, R., Navarete-Heredia, J.L., Beutel, R.G., 2005. 6.1. Lepiceridae. In: Beutel, R.G., Leschen, R.A.B. (Vol. Eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Archangelsky, M., Beutel, R.G., Komarek, A., 2005. 10. Hydrophiloidea. Introduction, Phylogeny. 10.1. Hydrophilidae. In: Beutel, R.G., Leschen, R.A.B. (Vol. Eds.), Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Arndt, E., Beutel, R.G., Will, K., 2005. 7.8. Carabidae. In: Beutel, R.G., Leschen, R.A.B. (Vol. Eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Balke, M., Ribera, I., Beutel, R.G., 2005. The systematic position of Aspidytidae, the diversification of Dytiscoidea (Coleoptera, Adephaga) and the phylogenetic signal of third codon positions. J. Zool. Syst. Evolut. Res. 43 (3), 223–242.

Beutel, R.G., & Friedrich, F., 2005. Comparative study of larvae of Tenebrionoidea (Cucujiformia, Coleoptera). Eur. J. Entomol. 102 (2), 241–164.

Beutel, R.G., & Weide, D., 2005. Cephalic anatomy of Zorotypus hubbardi (Hexapoda: Zoraptera): New evidence for a relationship with Acercaria. Zoomorph. 124, 121–136.

Beutel, R.G., 2005. 1. Systematic position and early evolution. 4. Morphology. 6. Myxophaga. Introduction, Phylogeny. 7.9. Rhysodidae. In: Beutel, R.G., Leschen, R.A.B. (Vol. Eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Beutel, R.G., Arce-Pérez, R., 2005. 6.3. Sphaeriusidae. In: Beutel, R.G., Leschen, R.A.B. (Vol. Eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Beutel, R.G., Arndt, E., 2005. 7.7. Trachypachidae. In: Beutel, R.G., Leschen, R.A.B. (Vol. Eds.), Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Beutel, R.G., Lawrence, J. F., 2005. 4. Morphology. In: Beutel, R.G.& Leschen, R.A.B. (Vol. Eds.), Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Beutel, R.G., Leschen, R.A.B. (Eds.) 2005. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). In: Band/Volume IV Arthropoda: Insecta Teilband / Part 38. Handbook of Zoology. Berlin, New York, De Gruyter.

Beutel, R.G., Leschen, R.A.B. (vol. eds), 2005. Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Beutel, R.G., Leschen, R.A.B., 2005. 8. Polyphaga. Introduction, Phylogeny. 9. Staphyliniformia. Introduction, Phylogeny. 18.10. Chelonariidae. In: Beutel, R.G., Leschen, R.A.B. (Vol. Eds.). Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Beutel, R.G., Leschen, R.A.B., 2005. Phylogenetic analysis of Staphyliniformia (Coleoptera) based on characters of larvae and adults. Syst. Entomol. 30, 510–548.

Beutel, R.G., Pohl, H., 2005. Head structures of males of Strepsiptera (Hexapoda) with emphasis on basal splitting events within the Order. J. Morph. 244, 1–14.

Beutel, R.G., Pohl, H., Hünefeld, F., 2005. Strepsipteran brains and effects of miniaturisation. Arthropod Str. Dev. 34, 301–313.

Beutel, R.G., Ribera, I., 2005. 7. Adephaga. Introduction, Phylogeny. In: Beutel, R.G., Leschen, R.A.B. (Vol. Eds.), Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Beutel, R.G., Roughley, R.E., 2005. 7.1. Gyrinidae. In: Beutel, R.G., Leschen, R.A.B. (Vol. Eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Beutel, R.G., Vanin, S.A., 2005. 6.2. Torridincolidae. In: Beutel, R.G., Leschen, R.A.B. (Vol. Eds.), Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Hünefeld, F., Beutel, R.G., 2005. The sperm pumps of Strepsiptera and Antliophora (Hexapoda). J. Zool. Syst. Evolut. Res. 43 (4), 297–306.

Jäch, M., Beutel, R.G., Delgado, J.A., Díaz, J.A., 2005. 11.1. Hydraenidae. In: Beutel, R.G., Leschen, R.A.B. (Vol. Eds.), Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Navarrete-Heredia, J.L., Cortés-Aguilar, J., Beutel, R.G., 2005. New findings on the enigmatic beetle family Lepiceridae (Coleoptera: Myxophaga). Entomol. Abh. 62 (2), 193–201.

Pohl, H., Beutel, R.G., 2005. The phylogeny of Strepsiptera (Hexapoda). Cladistics 21,1–47.

Pohl, H., Beutel, R.G., Kinzelbach, R., 2005. Protoxenidae fam. n. (Insecta, Strepsiptera) from Baltic amber – a ‘missing link’ in strepsipteran phylogeny. Zool. Scr. 31 (1), 123–134.

Vanin, S.A., Beutel, R.G., Arce-Pérez, R., 2005. 6.4. Hydroscaphidae. In: Beutel, R.G., Leschen, R.A.B (Vol. Eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Alarie, Y., Beutel, R.G., Watts, C.H.S., 2004. Larval morphology of three species of Hygrobiidae (Coleoptera: Adephaga: Dytiscoidea) with phylogenetic considerations. Eur. J. Entomol. 101, 293–311.

Anton, E., & Beutel, R.G., 2004. On the head morphology and systematic position of Helophorus (Coleoptera: Hydrophiloidea: Helophoridae). Zool. Anz. 242, 313–346.

Beutel, R.G., 2004. Aktuelle Entwicklungen in der Systematik der Hexapoda (Arthropoda). Entomol. heute 16, 21–50.

Beutel, R.G., Komarek, A., 2004. Comparative study of thoracic structures of adults of Hydrophiloidea and Histeroidea with phylogenetic implications (Coleoptera, Polyphaga). Org. Divers. Evol. 4, 1–34.

Korte, A., Ribera, I., Beutel, R.G., Bernhard, D., 2004. Interrelationships of Staphyliniform groups inferred from 18S and 28S rDNA sequences, with special emphasis on Hydrophiloidea (Coleoptera, Staphyliniformia). J. Zool. Syst. Evolut. Res. 42 (4), 281–288.

Leschen, R.A.B., Beutel, R.G., 2004. Ocellar atavism in Coleoptera: plesiomorphy or apomorphy? J. Zool. Syst. Evolut. Res. 42 (1), 63–69.

Pohl, H., Beutel, R.G., 2004. Fine structure of adhesive devices of Strepsiptera (Insecta). Arthropod Str. Dev. 33, 31–43.